1) Echolon 1 - The Honey Bee Swarm
Small, cheap and endurant VLO variants of UAV like the Scaneagle or Integrator fitted with various sensor packages will be able to penetrate an enemy airspace with a very low risk of detection. This swarm will be equipped with a combination of sensor packages consisting of IRST (Infrared Search and Track), RWR (Radar Warning Receiver) or/and AESA radars will ISR the area. The "hot" compontents of the GBAD will be geolocalized. Even opposing "cold" VLO fighters like the F-22 controlled by an AEW&C might get detected by the dispersed swarm of very compact VLO UAV when they try to close in with the screened AEW&C which is hanging back 150+ km. Some Bees might be used as comm. relays.
2) Echolon 2 - The Bumblebee Swarm
VLO UCAV like the Avenger C or X-45 are the "bomb trucks" and might also carry a limited number of AAM. Heavier, larger and faster they are more suscitable to detection but the fine ISR or the honey bees will allow them to increase their surviveability. SEAD and high-value targets their secundary. For self- or AEW&C defense they might use their AAM. BVR might be a feasible addition and could be offer on a relative fast VLO UCAV supported by the AEW&C rather impressive attack capabilities against an manned interceptor vectored for the AEW&C, the "honey pot" aka "bee queen".
3) Echolon 3 - The Bee queen
The AEW&C will coordinate the C4ISR and help to weave the picture togheter from a safe distance. A second AEW&C might be sometimes necessary as the workload of the Bee Quenn will be heavy and slow, endurant Bees and Bumblebees might require a great persistence of their Queen.
4) Echolon 4 - The Hornets
Modern manned mulit-role fighters will according to the mission guard the Bee queen and possibly the Bumblebees, create air superiority and/or deliver strikes with guided bombs and stand-off weapons.
5) Ground- or Seabased assets might deliver heavy long-ranged precision firepower and reduce the strain on the UAV's payload
6) Tankers will offer in the midterm air refueling to the "bumblebee" class of UCAV, while the "bee" class of ISR UAV will most likely have to rely on ground refueling.
BTW: The JSF would IMHO both be an excellent combination of an "Hornet" or and a "Bumblebee", earining them then name "Wasp" or perhaps "Aculeata" if we want to keep the somewhat corret biological taxonomy